I recommend buying the book "Force: Character Design from Life Drawing" by Mike Mattesi (which features John Watkiss’s artwork on many pages), as it contains the high-resolution version of what those PDFs show in low quality.

However, it is important to clarify that (the renowned British storyboard artist and concept artist) did not write a standard academic medical anatomy paper. Instead, he is famous for his dynamic figure drawing workshops and his book "Force: Dynamic Life Drawing for Animators" (though that book is by Mike Mattesi, Watkiss is a major contributor to the philosophy of gesture and force ).

The most useful paper (or PDF resource) that aligns with is typically a compiled set of his handout notes from his lectures at Sundance (formerly the Disney-led training program) or Gobelins .